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bear

 
Dictionary: bear2   (bâr) pronunciation
Bear
Bear
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n.
    1. Any of various usually omnivorous mammals of the family Ursidae that have a shaggy coat and a short tail and walk with the entire lower surface of the foot touching the ground.
    2. Any of various other animals, such as the koala, that resemble a true bear.
  1. A large, clumsy, or ill-mannered person.
    1. One, such as an investor, that sells securities or commodities in expectation of falling prices.
    2. A pessimist, especially regarding business conditions.
  2. Slang. Something that is difficult or unpleasant: The final exam was a bear.
  3. Slang. A highway patrol officer.
adj.
Characterized by falling prices: a bear market.

[Middle English bere, from Old English bera. Sense 3, probably from proverb To sell the bear's skin before catching the bear.]


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Generally massive, short-legged mammals constituting the family Ursidae. Bears are the most recently evolved carnivore, found in Europe, Asia, and the Americas. Closely related to the dog and the raccoon, most bears climb with ease and are strong swimmers. As a family, they are omnivores, but dietary preferences vary among species (the polar bear feeds mainly on seals, the spectacled bear on vegetation, etc.). Though they do not truly hibernate, bears often sleep fitfully through much of the winter. They live 15 – 30 years in the wild but much longer in captivity. They have been hunted as trophies, for hides, and for food. See also black bear; brown bear; sun bear.

For more information on bear, visit Britannica.com.

(Ursidae)

Class: Mammalia

Order: Carnivora

Suborder: Fissipedia

Family: Ursidae

Thumbnail description
Medium to large, stocky mammals with fur that may be black, brown, reddish, or white

Size
Adults average 4–5 ft (1.2–1.5 m) and 60–150 lbs (27–70 kg) in body length for sun bears to 8–9 ft (2.4–2.7 m) and 900–1,300 lbs (400–590 kg) for polar bears

Number of genera, species
3–6 genera; 8 (or more) species

Habitat
Wide ranging, including forests, rainforests, tundra, deserts, and swamps

Conservation status
Endangered: 1 species; Vulnerable: 3 species; Lower Risk/Conservation Dependent: 1 species; Data Deficient: 1 species

Distribution
On every continent except Africa and Antarctica, but mainly in the Northern Hemisphere

Evolution and systematics

Although this family has a small number of genera and species, it still has a good share of controversy when it comes to classification. For example, some systematists over the years have placed the giant panda in a subfamily of Ursidae, as it is in this chapter, or in its own family, called Ailuropodidae. The Malayan sun bear, sloth bear, and polar bear are often grouped under the Ursus genus, but sometimes fall under the genera Helarctos, Melursus, and Thalarctos, respectively. Subspecies of the brown bear (U. arctos) are often listed as separate species, including the Alaskan brown bear (U. middendorffi) and the grizzly bear (U. horribilis). In addition, the red panda (Ailurus fulgens) was once listed with the ursids, but is now considered a to be a member of its own family, the Ailuridae, or a sub-family of the Procyonidae, which includes the raccoons.

This chapter uses the following classification:

  • giant panda, Ailuropoda melanoleuca
  • Malayan sun bear, Helarctos malayanus
  • sloth bear, Melursus ursinus
  • spectacled bear, Tremarctos ornatus
  • American black bear, Ursus americanus
  • brown bear, U. arctos
  • polar bear, U. maritimus
  • Asiatic black bear, U. thibetanus

The family Ursidae is believed to have originated in Asia, and is closely related to the canids (dogs and relatives), procyonids (raccoons and relatives), and ailurids (lesser panda). The giant panda is considered to be the most primitive of the bears. Various evolutionary studies have attempted to determine the relationships of the other bears. Fossil studies seem to indicate that the spectacled bear, which is in the subfamily Tremarctinae, diverged from the remaining bears, which are in the subfamily Ursinae. The fossil record also points to a very close relationship between the Asiatic black and American black bears, and places brown and polar bears close to them evolutionarily. Other studies using mitochondrial DNA and cytochrone-b sequence data have provided clarification, and sometimes challenged, previous conclusions. For example, mtDNA data have indicated that polar bears and spectacled bears are very closely related, and diverged from the ursinids about 2 million years ago. Cytochrome-b data appear to show that the sun bear and American black bear are sister taxa, and are somewhat separated from the Asiatic black bear.

Physical characteristics

Bears are medium to large, powerful mammals with rather short tails and plantigrade feet on stocky limbs. Many bears are dark brown to black, but the fur color is often variable within the species and sometimes even among siblings. Among the American black bear, for instance, black, brown, reddish, and even whitish individuals exist. Some species have distinctive white patches or lines on the face, throat, and/or chest. An example is the spectacled bear, which has whitish rings around its eyes. The polar bear is the only species that consists of all white-furred individuals, although the skin is black to make the best use of the heat from the arctic sun, and the individual outer (or guard) hairs are actually clear rather than white. Some species, such as the brown bear, have longer hair on the shoulders that forms a mane. A few, like the sloth bear, have long fur over much of their bodies.

Their heads are rather large, particularly in some species like the panda, and they have small, forward-facing eyes, and noticeable but usually modest round ears. Their teeth include premolars and molars designed for crushing, and long canines, which together assist their omnivorous diet. The giant panda and spectacled bear have flattened molars suited to their strongly herbivorous diets. The sloth bear, which is particularly fond of termites, has no incisors in its upper jaw. The gap, combined with protrusible and naked lips, allows the bear to suck up the insects. Sun bears have especially long tongues to assist them in attaining honey, a frequent item in their diet.

Bear claws, which are non-retractile, differ in length in separate species. In the brown bear, for example, the light-colored claws stand out from the typically brown fur, and range from 2–4 in (5–10 cm) long. The Asiatic black bear, on the other hand, has comparatively short claws, typically measuring less than 2 in (4–5 cm) in length.

In overall size, bears have a fairly wide range. In all species, males are larger than females. The smallest ursid is the Malayan sun bear, with a body length of 4–5 ft (1.2–1.5 m), and a typical male weight of 60–150 lb (27–70 kg). Polar and brown bears are at the opposite end of the spectrum, with male polar bears averaging 8–9 ft (2.4–2.7 m) in body length and 900–1,300 lb (400–590 kg), and male brown bears 5–8 ft (1.5–2.4 m) and about 350–850 lb (160–385 kg).

Distribution

Bears have a wide distribution in the Northern Hemisphere. Here, the most widely distributed species include the polar bears, which inhabit the circumpolar ice pack; the brown bears, which live throughout northern North American and north to north-central Eurasia; and the American black bear, which stretches from northern Mexico well into Canada. Both the sun bear and sloth bear reside in Southeast Asia. The range of the Asiatic black bear is somewhat larger, extending from Afghanistan to southeastern Russia. The giant panda has the smallest range of all bears, with six small populations known from the Tibetan plateau in southwestern China. The spectacled bear, the only ursid that inhabits in the Southern Hemisphere, lives around the Andes in South America, including sites in Bolivia, Colombia, Ecuador, Peru, and Venezuela.

Habitat

Bears' habitats vary from species to species. The polar bear thrives on the arctic ice pack, a sharp contrast to the tropical rainforests of southeast Asia, where the Malayan sun bear resides. The American black bear's habitat spreads from the woods in western U.S. mountains to wetlands in southeastern states, and to the northern tundra in Canada. In contrast, the shaggy-looking sloth bear opts for grasslands and dry forests from lowlands in India to the foothills of the Himalayas.

The brown bear, also known in parts of North America as the grizzly or kodiak, ranges from thickly forested areas into grasslands and tundra in the Northern Hemisphere, while the spectacled bear prefers lush mountain forests in South America. The giant panda lives in the bamboo forests of China, and the Asiatic black bear in primarily moist forests throughout southern Asia.

Home ranges for bears also vary. Pandas keep to about 2–3 mi2, but brown bears are known to range over 800–1,000 mi2 if the habitat is poor and food is scarce.

Behavior

Little is known about the behavior in the wild of half of the ursid species, mostly due to their remote geographical distribution. These include the spectacled bear, Asiatic black bear, sun bear, and sloth bear. In general, however, bears overall are solitary animals except during mating season or in mother-cub groupings. Occasionally among brown bears, siblings will stay near one another for a year or two after they leave their mother. Although additional research is needed for substantiation, some reports indicate that sloth bears may form social units, and that male sun bears may remain with the mother after she gives birth.

Bears generally maintain home ranges, with the males' ranges frequently overlapping with those of the females. Black bears mark their territories with scent markings or long scratches clawed into trees. Unusually, male panda bears sometimes do their scent marking while standing on their hands. In black bears and several other species, the ranges of male bears may also overlap, but since the ranges are often very large and bears rarely see one another, the overlaps present little opportunity for territorial conflicts. Even when bears come together at one feeding site, such as brown or black bears at a salmon stream, individual bears maintain their personal space and share the resource. When bears approach one another too closely, temporary dominance hierarchies may form, with the largest males mounting short-lived aggressive displays, including growls, and occasional charges to maintain a small feeding territory. During breeding season, males generally compete for females, but the male-female bonds typically only last one or two weeks.

With their large, plantigrade feet and stout limbs, bears are often pictured as lumbering animals that always move slowly and deliberately. They can, however, move very quickly when necessary. Black bears, for example, can run at speeds of 30 mph (50 kph), and polar bears are fast enough to catch caribou on the Arctic tundra. Even the somewhat awkward-moving sloth bear can outrun a human over short distances. Most bears are also accomplished tree climbers. The sun bear has perfected climbing, quickly scaling trees in search of honey and other food items, and even fashioning resting/feeding platforms out of broken branches high up in the trees. Polar bears and adult brown bears (with the exception of some populations in Europe) do not climb, but both are good swimmers. Other bears, like the Asiatic black bear, can also swim. With their large and slightly webbed front feet, polar bears are particularly adept swimmers and divers, and reportedly are able to swim across open-water expanses of up to 100 mi (65 km).

Ursids tend to be crepuscular (mainly active at dawn and dusk) or nocturnal animals, although some extend their active periods into the daytime. Polar bears are an example. While they are most active at night and at dawn, they are frequently seen hunting during the day.

Although ursids do not technically hibernate, many cooler-climate bears do enter winter dormancy, during which the respiratory and heart rates drop, but the body temperature dips only slightly. In the black bear, for instance, their body temperature drops from about 100°F (38°C) to 88–93°F (31–34°C). The Asiatic black bear is an exception: Its body temperature declines precipitously to just 37–45°F (3–7°C). It is during the winter dormancy that female ursids give birth. As she sleeps, the young suckle and grow. Among the cooler-climate species, both males and females become dormant, except in the polar bears, where only pregnant females enter winter sleep. During the winter, bears are capable of awakening, and occasionally leave their winter dens, which may be burrows, hollow logs, or tunnels in the snow and ice. Warm-climate bears, including sun, sloth, and spectacled bears, do not enter winter sleep. Although most brown bears and Asiatic black bears "hibernate," those from warmer climates frequently skip winter dormancy and remain active all year.

Feeding ecology and diet

Bears are omnivores, often eating whatever is available. The polar bear tends heavily toward a carnivorous diet, existing primarily on ringed seals (Phoca hispida), although it will eat berries and vegetation in the summer. Hunting is accomplished either by ambush or active stalking. In the former, the polar bear will simply wait at an ice hole for a seal to surface, then overpower it with one whack of its mighty paw. It reverts to stalking either on land or in the water if it happens to see a seal on the ice or another animal, such as an elk, in the open. With the polar bear's cryptic coloration, it can approach closely enough on land to give chase and sometimes overtake the animal.

Other ursids tend to prefer a greater amount of vegetation than the polar bear, eating fruits, tender stems, and roots most of the time, and supplementing the diet with insects, fish, an occasional small mammal, or carrion. The larger bears, like the brown bear, will sporadically hunt moose, or elk (Alces alces), and other ungulates. The giant panda, on the other hand, is almost exclusively an herbivore, eating little but bamboo leaves, stems and shoots. Sloth bears are unusual in their strong reliance on termites for food, although other bears also eat termites to some extent. Sloth bears, along with the sun bear in particular, are also fond of honey. In both cases, the bears use their claws to rip open termite and bees nests, and get at the reward.

Reproductive biology

Bears' mating systems vary by species; some, such as the spectacled bear, are monogamous. Others, such as the polar bear, are polygamous. Most bears mate in the spring or summer, but the fertilized eggs do not implant in the uterus and begin developing until fall. After this so-called delayed implantation, the eggs begin to develop and the females give birth in the winter. Some species, including the sloth bear, apparently mate year-round in especially warm climates, but due to delayed implantation, all give birth in the winter. Sun bears appear to have delayed implantation, but individuals in zoos have given birth at different times of the year.

Cubs are born small, naked and blind, having developed in the womb for only two to three months. Birth weight ranges from about 11 oz (325 g) in sun bears to 21 oz (600 g) in brown and polar bears. In most cases, females give birth from one to five cubs, although two is the most common litter size among ursids. Panda mothers generally rear only one cub, regardless of the litter size, and the others die. Among cooler-climate bears, the young are born while the mother is in winter dormancy. In warmer-climate species, such as the sun bear, the mother chooses a concealed site, perhaps under branches or thick vegetation, to make a nest for the cubs. Cubs are generally weaned within the first two to five months (pandas wean at about nine months), but remain with the family unit for two to four years, during which the cubs learn to find their own food and hunt while under the protective eye of their mother.

Sexual maturity generally occurs from four to seven years old, but the timing varies among species.

Conservation status

Only one ursid, the panda bear, has been listed as Endangered by the IUCN. Habitat destruction is a major reason for this species' decline. A recent study indicated that not only human population magnitude, but the increasing number of households as family units decrease in size, have contributed to intensifying habitat destruction, particularly for this bear. As the number of households have risen, deforestation and fragmentation of panda habitat have accelerated. Estimates place the total number of pandas in the wild at below 1,000.

Other bears have also experienced decreased suitable habitat and habitat fragmentation. The brown bear, for instance, is now found in only 2% of its former range within the continental United States. The lessened range is blamed in part on habitat destruction and fragmentation. In addition, individual populations of various species have experienced declines, even if the overall species numbers are relatively high. The American black bear is an example. Several subspecies, including Ursus americanus floridanus, are considered threatened locally. To counteract the declines, various hunting bans or regulations, habitat preservation programs, and educational efforts are under way worldwide.

The sloth bear, spectacled bear, and Asiatic black bear are listed as Vulnerable; the polar bear is Lower Risk/Conservation Dependent; and the Malayan sun bear is listed as Data Deficient by the IUCN.

Significance to humans

Various bears are hunted for meat, fur, and trophy mounts. In addition, body parts, such as the gall bladder of sun and American black bears, are also harvested for medicinal purposes, particularly in China. Bears have also become important as attractions at zoos.

While many bears are assumed to be dangerous to humans, bear attacks are few and fatalities are rare.

Species accounts

American black bear
Brown bear
Giant panda
Polar bear
Spectacled bear

Resources

Books:

Craighead, L. Bears of the World. Blaine, WA: Voyager Press, 2000.

Heldmaier, G. and M. Klingenspor, eds. Life in the Cold: The 11th International Hibernation Symposium. New York: Springer-Verlag, 2000.

Kays, R., and D. Wilson. Mammals of North America (Princeton Field Guides). Princeton: Princeton University Press, 2002.

Wilson, D., and Reeder, D. Mammal Species of the World, a Taxonomic and Geographic Reference. 2nd ed. Washington, DC: Smithsonian Institution Press, 1993.

Periodicals:

Liu, J., G. C. Daily, P. R. Ehrlich, and G. W. Luck. "Effects of household dynamics on resource consumption and biodiversity." Nature 421 (Jan. 12, 2003): 530–533.

Loucks, C., et al. "Giant pandas in a changing landscape." Science 294 (Nov. 16 2001): 1465

Milius, Susan. "The lives of pandas." Science News 159 (Jan. 27, 2001): 61–3.

Mills, J. "Milking the bear trade (for their bile; sidebar with illustrations and data on bears throughout the world)." International Wildlife 22 (1992): 38–45.

Slattery, J., and S. O'Brien. "Molecular phylogeny of the red panda (Ailurus fulgens)." The Journal of Heredity 86 (November/December 1995): 413–22.

Tyson, P. "Secrets of Hibernation,"

Zhang, Y-P., Ryder, and A. Oliver. "Mitochondrial DNA sequence evolution in the Arctoidea." Proceedings of the National Academy of Sciences of the United States of America 90 (1993): 9557–61.

Organizations

American Bear Association. P.O. Box 77, Orr, MN 55771 United States. Phone: (218) 757-0172. E-mail: admin@americanbear.org Web site:

Great Bear Foundation. P.O. Box 9383, Missoula, MT 59807 United States. Phone: (406) 829-9378. Fax: (406) 829-9379. E-mail: gbf@greatbear.org Web site:

IUCN/SSC Bear Specialist Group. Harry V. Reynolds, Alaska Dept. of Fish and Game-Fairbanks, 1300 College Rd., Fairbanks, AK 99701 United States. Phone: (907) 459-7238. Fax: (907) 459-9723. E-mail: harry_reynolds@fishgame.state.ak.us Web site:

Ursus International Conservation Institute. P.O. Box 832, Pincher Creek, Alberta T0K 1W0 Canada. E-mail: info@ursusinternational.org Web site:

Other

Animal Diversity Web. University of Michigan Museum of Zoology.

The Bear Den.

Bears.org.

Hilton-Taylor, C., (comp.) 2000 IUCN Red List of Threatened Species. IUCN/SSC: Gland, Switzerland and Cambridge, UK, 2000.

Nowak, R. "Black, Brown, Polar, Sun, and Sloth Bears." In Walker's Mammals of the World Online. The Johns Hopkins University Press, 1995.

National Science Foundation. Press release NSFPR 03-06. Jan. 12, 2003. "Researchers Tie Worldwide Biodiversity Threats to Growth in Households: Pandas in China face encroachment, as do other species in global hotspots."

Smithsonian National Museum of Natural History.

[Article by: Leslie Ann Mertz, PhD]

Several species of this large mammal, family Ursidae, were common in western and northern Europe until modern times so that the bear was, along with the boar, one of the most ferocious native beasts a person might encounter. Not surprisingly, then, the bear figured often in the Celtic imagination. Bears are found on Gaulish coins and statuettes. The bear-goddess Artio, also known as Andarta, was venerated at what is now Berne (‘bear city’), Switzerland. Another bear divinity was Matus or Matunus, venerated at Risingham, north of Hadrian's Wall. Names deriving from native forms for ‘bear’ are cited often in Welsh and Irish genealogies. The Welsh for bear is arth or arthen; cf. Breton arzh, giving rise to the Gaelic personal name Art and possibly also Arthur. Archaic Old Irish for bear is math; Old Irish and MidIr. mathgamain; Modern Irish mathghamhain; Scottish Gaelic mathghamhainn. See also BEITHIR.

 
bear, large mammal of the family Ursidae in the order Carnivora, found almost exclusively in the Northern Hemisphere. Bears have large heads, bulky bodies, massive hindquarters, short, powerful limbs, very short tails, and coarse, thick fur. They walk on the entire sole of the foot and normally move with a slow, ambling gait. However, they are capable of moving with great speed when necessary and some achieve bursts of 35 mi (56 km) per hour. Most bears can climb trees and swim well. They stand on the hind feet to reach objects with their paws. They have large, strong, non-retractile claws, used for catching prey and for digging. Their teeth are adapted to grinding as well as tearing. Nearly all species are omnivorous, feeding on fruits, roots and other plant matter, honey, carrion, insects, fish, and small mammals.

Adult bears are solitary except during the mating season. Groups may feed together where quantities of food are available, but there is little social contact. In cold climates bears sleep through most of the winter in individual dens made in caves or holes in the ground. This sleep is not a true hibernation, as the bear's metabolism remains in a normal state and it may wake and emerge during warm spells. The young, usually twins, are born during winter in a very immature state. Cubs stay with their mothers for about a year, and females usually mate only every other year. Bears are not generally subject to predation, unless they are in a weakened condition. A bear is a formidable adversary and may attack a human if it is injured or startled.

Types of Bears

The brown bear of Eurasia, Ursus arctos, is extinct in much of Western Europe, but small numbers survive in some wooded sections of that region and larger numbers in Russia and N Asia. The Russian variety was the bear most often trained to dance and box in circuses and shows in the past.

The Asian black bear, or moon bear, Selenarctos thibetanus, is found in forests from central Asia and the Himalayas to Japan. The sun bear, Helarctos malayanus, is found in tropical forests of SE Asia. Smallest of the bears, it is about 4 ft (120 cm) long and weighs about 100 lb (45 kg). It spends much time in trees and is fond of honey; it is sometimes called honey bear (a name also applied to the kinkajou). The sloth bear, Melursus ursinus, is a medium-sized bear of the forests of S India and Sri Lanka.

The North American brown bears, including the Kodiak bear and grizzly bear, are regarded by many authorities as varieties of U. arctos. Brown bears are dish-faced; i.e., their muzzles curve upward in profile. Their shoulders are humped. They range in color from yellow-brown to nearly black, with much color variation among different varieties, local populations, and individuals. Most varieties do not climb well. The Kodiak bear, or big brown bear, is the largest living member of the Carnivora, sometimes reaching a length of 9 ft (2.7 m), a shoulder height of 41/2 ft (140 cm), and a weight of over 1,600 lb (730 kg). It is found along the south coast of Alaska and, like the Siberian brown bear, eats large numbers of salmon during salmon runs.

The most widespread and numerous North American bear is the so-called black bear, U. americanus, found in Alaska, Canada, the Great Lakes region, mountainous areas of the United States, and on the Gulf Coast. American black bears range in color from light brown to black; in northern regions there are gray and nearly white forms. Their muzzles are always cinnamon brown and are straight in profile. They are further distinguished from brown bears by their smaller size and by their hindquarters, which are higher than their shoulders. Males are usually about 6 ft (190 cm) long and weigh about 500 lb (230 kg).

The polar bear, Thalarctos maritimus, is an almost exclusively carnivorous species of the Arctic. The only bear of the Southern Hemisphere is the spectacled bear, Tremarctos ornatus, of the Andes Mts.; it is so called from the light-colored circles around its eyes. Recent evidence suggests also including the giant panda, Ailuropoda melanoleuca as a bear.

Classification

Bears are classified in the phylum Chordata, subphylum Vertebrata, class Mammalia, order Carnivora, family Ursidae.

Bibliography

See R. Perry, Bears (1970).


Wikipedia: Bear
Top
Bears
Fossil range: 38–0 Ma
Late Eocene - Recent
American Black Bear, Ursus americanus
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Caniformia
Family: Ursidae
G. Fischer de Waldheim, 1817
Genera

Bears are mammals of the family Ursidae. Bears are classified as caniforms, or doglike carnivorans, with the pinnipeds being their closest living relatives. Although there are only eight living species of bear, they are widespread, appearing in a wide variety of habitats throughout the Northern Hemisphere and partially in the Southern Hemisphere. Bears are found in the continents of North America, South America, Europe, and Asia.

Common characteristics of modern bears include a large body with stocky legs, a long snout, shaggy hair, plantigrade paws with five nonretractile claws, and a short tail. While the polar bear is mostly carnivorous and the giant panda feeds almost entirely on bamboo, the remaining six species are omnivorous, with largely varied diets including both plants and animals.

With the exceptions of courting individuals and mothers with their young, bears are typically solitary animals. They are generally diurnal, but may be active during the night (nocturnal) or twilight (crepuscular), particularly around humans. Bears are aided by an excellent sense of smell, and despite their heavy build and awkward gait, they can run quickly and are adept climbers and swimmers. In autumn some bear species forage large amounts of fermented fruits which affects their behaviour.[1] Bears use shelters such as caves and burrows as their dens, which are occupied by most species during the winter for a long period of sleep similar to hibernation.

Bears have been hunted since prehistoric times for their meat and fur. To this day, they play a prominent role in the arts, mythology, and other cultural aspects of various human societies. In modern times, the bear's existence has been pressured through the encroachment on its habitats and the illegal trade of bears and bear parts, including the Asian bile bear market. The IUCN lists six bear species as vulnerable or endangered, and even least concern species such as the brown bear are at risk of extirpation in certain countries. The poaching and international trade of these most threatened populations is prohibited, but still ongoing.

Contents

Evolutionary relationships

Fossil of Cave bear (Ursus spelaeus)

The Ursidae family belongs to the order Carnivora and is one of nine families in the suborder Caniformia, or "doglike" carnivorans. Bears' closest living relatives are the pinnipeds, a clade of three families: Odobenidae (the walrus), Otariidae (fur seals and sea lions), and Phocidae (true or earless seals). Bears comprise eight species in three subfamilies: Ailuropodinae (monotypic with the giant panda), Tremarctinae (monotypic with the Spectacled Bear), and Ursinae (containing six species divided into one to three genera, depending upon authority).

The origins of Ursidae can be traced back to the very small and graceful Parictis that had a skull only 7 cm (3 in) long. Parictis first occur in North America in the Late Eocene (ca. 38 million years ago), but this genus did not appear in Eurasia and Africa until the Miocene.[2] The raccoon-sized, dog-like Cephalogale, however, is widely regarded as the most primitive ursid and is ideally suited as a representative basal taxon for the family. Cephalogale first appeared during the middle Oligocene and early Miocene (approximately 20–30 million years ago) in Europe. Cephalogale gave rise to a lineage of early bears of the genus Ursavus. This genus radiated in Asia and ultimately gave rise to the first true bears (genus Ursus) in Europe, 5 million years ago. Even among its primitive species, such as C. minor, it exhibits typical ursid synapomorphic dentition such as posteriorly oriented M2 postprotocrista molars, elongated m2 molars, and a reduction of the premolars. Living members of the ursids are morphologically well defined by their hypocarnivorous (non-strictly meat-eating) dentitions, but fossil ursids include hypercarnivorous (strictly meat-eating) taxa, although they never achieved the extreme hypercarnivory seen in mustelids. Cephalogale was a mesocarnivore (intermediate meat-eater).[3] Other extinct bear genera include Arctodus, Agriarctos, Plionarctos and Indarctos.

It is uncertain whether ursids were in Asia during the late Eocene, although there is some suggestion that a limited immigration from Asia may have produced Parictis in North America due to the major sea level lowstand at ca. 37 Ma, but no Parictis fossils have yet to be found in East Asia. Ursids did, however, become very diversified in Asia later during the Oligocene. Four genera representing two subfamilies (Amphicynodontinae and Hemicyoninae) have been discovered in the Oligocene of Asia: Amphicticeps, Amphicynodon, Pachycynodon, and Cephalogale. Amphicticeps is endemic from Asia and the other three genera are common to both Asia and Europe. This indicates migration of ursids between Asia and Europe during the Oligocene and migration of several taxa from Asia to North America likely occurred later during the late Oligocene or early Miocene. Although Amphicticeps is morphologically closely related to Allocyon, and also to Kolponomos of North America, no single genus of the Ursidae from this time period is known to be common to both Eurasia and North America. Cephalogale, however, do appear in North America in the early Miocene. It is interesting to note that rodents, such as Haplomys and Pseudotheridomys (late Oligocene) and Plesiosminthus and Palaeocastor (early Miocene), are common to both Asia and North America and this indicates that faunal exchange did occur between Asia and North America during the late Oligocene to early Miocene. Ursid migration from Asia to North America would therefore have also been very likely to occur during this time.[4] In the late Neogene three major carnivoran migrations that definitely included ursids are recognized between Eurasia and North America. The first (probably 21–18 Ma) was waves of intermittent dispersals including Amphicynodon, Cephalogale and Ursavus. The second migration occurred about 7–8 Ma and included Agriotherium (probably a hemicyonid – this was unusual among ursoids in that it also colonised sub-Saharan Africa). The third wave took place in the early Pliocene 4 Ma, consisting of Ursus.[5]

The giant panda's taxonomy has long been debated. Its original classification by Armand David in 1869 was within the bear genus Ursus, but in 1870 it was reclassified by Alphonse Milne-Edwards to the raccoon family.[6] In recent studies, the majority of DNA analyses suggest that the giant panda has a much closer relationship to other bears and should be considered a member of the family Ursidae.[7] The status of the red panda remains uncertain, but many experts, including Wilson and Reeder, classify it as a member of the bear family. Others place it with the raccoons in Procyonidae or in its own family, the Ailuridae. Multiple similarities between the two pandas, including the presence of false thumbs, are thought to represent convergent evolution for feeding primarily on bamboo.

There is also evidence that, unlike their neighbors elsewhere, the brown bears of Alaska's ABC islands are more closely related to polar bears than they are to other brown bears in the world. Researchers Gerald Shields and Sandra Talbot of the University of Alaska Fairbanks Institute of Arctic Biology studied the DNA of several samples of the species and found that their DNA is different from that of other brown bears. The researchers discovered that their DNA was unique compared to brown bears anywhere else in the world. The discovery has shown that while all other brown bears share a brown bear as their closest relative, those of Alaska's ABC Islands differ and share their closest relation with the polar bear.[8] There is also the very rare Tibetan Blue Bear, which is a type of brown bear. This animal has never been photographed.

Koalas are often referred to as bears due to their appearance; they are not bears, however, but marsupials.

Classification

Brown Bear Ursus arctos, at the Moscow Zoo
Asian Black Bear Ursus thibetanus, at the Wrocław Zoo, Poland
Sun Bear Helarctos malayanus, at the Columbus Zoo
Giant Panda Ailuropoda melanoleuca, "Tian Tian"

The genera Melursus and Helarctos are sometimes also included in Ursus. The Asiatic black bear and the polar bear used to be placed in their own genera, Selenarctos and Thalarctos which are now placed at subgenus rank.

A number of hybrids have been bred between American black, brown, and polar bears (see Ursid hybrids).

Biology

Morphology

Despite being quadrupeds, bears can stand and sit similarly to humans.
Unlike other carnivora, bears have plantigrade hind feet

Bears are generally bulky and robust animals with relatively short legs. Bears are sexually dimorphic with regard to size, with the males being larger. Larger species tend to show increased levels of sexual dimorphism in comparison to smaller species, and where a species varies in size across its distribution individuals from larger sized areas tend also to vary more. Bears are the most massive terrestrial members of the order Carnivora, with some Polar Bears and Brown Bears weighing over 750 kilograms (1,700 lb). As to which species is the largest may depend on whether the assessment is based on which species has the largest individuals (brown bears) or on the largest average size (polar bears). The smallest bears are the Sun Bears of Asia, which weigh an average of 65 kilograms (140 lb) for the males and 45 kilograms (99 lb) for the females.[9]

Unlike other land carnivorans, bears are plantigrade. They distribute their weight toward the hind feet which makes then look lumbering when they walk. They are still quite fast with the brown bear reaching 30 miles per hour (48 km/h) although they are still slower than felines and canines. Bears can stand on their hind feet and sit up straight with remarkable balance. Bears have non-retractable claws which are used for digging, climbing, tearing and catching prey. Their ears are rounded.

Bears have an excellent sense of smell, a better sense of smell in fact that the dogs (Canidae), or possibly any mammal. This sense of smell is used for signalling between bears (either to warn off rivals or detect mates) and for finding food. Smell is the principal sense used by bears to find most of their diet.[9]

Dentition

Unlike most other members of the Carnivora, bears have relatively undeveloped carnassial teeth, and their teeth are adapted for a diet that includes a significant amount of vegetable matter. The canine teeth are large, and the molar teeth flat and crushing. There is considerable variation in dental formula even within a given species. It has been suggested that this indicates bears are still in the process of evolving from a carnivorous to a predominantly herbivorous diet. Polar bears appear to have secondarily re-evolved fully functional carnassials, as their diet has switched back towards carnivory.[10] The dental formula for living bears is: Upper: 3.1.2-4.2 / Lower: 3.1.2-4.3

Distribution and habitat

The bears are mostly found in the northern hemisphere, with a single species, the Andean Bear, occurring in South America. The Atlas Bear, a subspecies of the Brown Bear, was the only bear native to Africa. It was distributed in North Africa from Morocco to Libya, but has been extinct since around the 1870s. All the other species are found in North America, Asia and Europe. The most widespread species is the Brown Bear, which occurs from Western Europe eastwards through Asia to the western areas of North America. The American Black Bear is restricted to North America, and the Polar bear is restricted to the Arctic Sea. All the remaining species are Asian.[9]

With the exception of the Polar Bear the bears are mostly forest species. Some species, particularly the Brown Bear, may inhabit or seasonally use other areas such as alpine scrub or tundra.

Behaviour

While many people think that bears are nocturnal, they are in fact generally diurnal, active for the most part during the day. The belief that they are nocturnal apparently comes from the habits of bears that live near humans which engage in some activities, such as raiding trash cans or crops, are nocturnal in order to avoid humans. The sloth bear of Asia is the most nocturnal of the bears, but this varies by individual and females with cubs are often diurnal in order to avoid competition with males and nocturnal predators.[9] Bears are overwhelming solitary and are considered to be the most asocial of all the Carnivora. Liaisons between breeding bears are brief, and the only times bears are encountered in small groups are mothers with young or occasional seasonal bounties of rich food (such as salmon runs).[9]

Vocalizations

Bears produce a variety of vocalizations such as:

  • Moaning: produced mostly as mild warnings to potential threats or in fear.
  • Barking: produced during times of alarm, excitement or to give away the animal's position.
  • Huffing: made during courtship or between mother and cubs to warn of danger.
  • Growling: produced as strong warnings to potential threats or in anger.
  • Roaring: used much for the same reasons as growls and also to proclaim territory and for intimidation. Made by all bears expect for the sun bear.

Diet and interspecific interactions

Brown Bears make use of infrequent but predictable salmon runs in order to feed

Their carnivorous reputation non-withstanding, most bears have adopted a diet of more plant than animal matter and are completely opportunistic omnivores. Some bears will climb trees in order to obtain mast (edible vegatative or reproductive parts such as acorns), smaller species which are more able to climb include a greater amount of this in their diet.[11] Such masts can be very important to the diet of these species, and mast failures may result in long range movements by bears looking for alternate sources of food.[12] One exception is the Polar Bear, which has adopted a diet mainly of marine mammals to survive in the Arctic. The other exception is the Giant Panda which has adopted a diet mainly of bamboo. Stable isotope analysis of the extinct Giant Short-faced Bear (Arctodus simus) shows that it was also an exclusive meat eater, probably a scavenger.[13] The Sloth Bear, though not as specialized as the previous two species, has lost several front teeth usually seen in bears and developed a long, suctioning tongue in order to feed on the ants, termites and other burrowing insects that they favour. At certain times of the year these insects can make up 90% of their diet.[14] All bears will feed on any food source that becomes available, and the nature of that varies seasonally. A study of Asiatic Black Bears in Taiwan found that they would consume large numbers of acorns when they were most common, and switch to ungulates in other times of the year.[15]

When taking warm-blooded animals, bears will typically take small or young animals, as they are easier to catch. Although (besides Polar Bears) both species of black bear and the Brown Bear can sometimes take large prey, such as ungulates.[15][16] Often, bears will feed on other large animals when they encounter a carcass, whether or not the carcass is claimed by or is the kill of another predator. This competition is the main source of interspecies conflict. Bears are typically the apex predators in their range due to their size and power, and can defend a carcass against nearly all comers. Mother bears also can usually defend their cubs against other predators. The Tiger is the only known predator known to regularly prey on adult bears, including Sloth Bears, Asiatic Black Bears, Giant Pandas, Sun Bears and small Brown Bears.

Breeding

Bear cubs, like this American Black Bear, are sometimes killed by males

The age at which bears reach sexual maturity is highly variable, both between and within species. Sexual maturity is dependent on body condition, which is in turn dependent upon the food supply available to the growing individual. In the females of smaller species may have young in as little as two years, whereas the larger species may not rear young until they are four or even nine years old. First breeding may be even later in males, where competition for mates may leave younger males without access to females.[9]

The bear's courtship period is very brief. Bears in northern climates reproduce seasonally, usually after a period of inactivity similar to hibernation, although tropical species breed all year round. Cubs are born toothless, blind, and bald. The cubs of brown bears, usually born in litters of 1–3, will typically stay with the mother for two full seasons. They feed on their mother's milk through the duration of their relationship with their mother, although as the cubs continue to grow, nursing becomes less frequent and cubs learn to begin hunting with the mother. They will remain with the mother for approximately three years, until she enters the next cycle of estrus and drives the cubs off. Bears will reach sexual maturity in five to seven years. Male bears, especially Polar and Brown Bears, will kill and sometimes devour cubs born to another father in order to induce a female to breed again. Female bears are often successful in driving off males in protection of their cubs, despite being rather smaller.

Winter dormancy

Cub polar bear is nursing 2.OGG
Polar bear mother is nursing her cub

Many bears of northern regions are assumed to hibernate in the winter. While many bear species do go into a physiological state often colloquially called "hibernation" or "winter sleep", it is not true hibernation. In true hibernators, body temperatures drop to near ambient and heart rate slows drastically, but the animals periodically rouse themselves to urinate or defecate and to eat from stored food. The body temperature of bears, on the other hand, drops only a few degrees from normal and heart rate slows only slightly. They normally do not wake during this "hibernation", and therefore do not eat, drink, urinate or defecate the entire period. Higher body heat and being easily roused may be adaptations, because females give birth to their cubs during this winter sleep.

Relationship with humans

Polar bear at Wapusk National Park, Canada

Some species, such as the polar bear, American black bear, Sloth Bear and the brown bear, are dangerous to humans, especially in areas where they have become used to people. All bears are physically powerful and are likely capable of fatally attacking a person, but they, for the most part, are shy, easily frightened and will avoid humans. Injuries caused by bears are rare, but are often widely reported.[17] The danger that bears pose is often vastly exaggerated, in part by the human imagination. However, when a mother feels her cubs are threatened, she will behave ferociously. It is recommended to give all bears a wide berth because they are behaviorally unpredictable.

Bears may also come into conflict with humans where they raid crops or attack livestock.[18][19] These problems may be the work of only a few bears but create a climate of conflict as farmers and ranchers may perceive all losses as due to bears and advocate the preventative removal of all bears.[19] Mitigation methods may be used to reduce bear damage to crops, and reduce local antipathy towards bears.[18]

Laws have been passed in many areas of the world to protect bears from hunters habitat destruction. Public perception of bears is often very positive, as people identify with bears due to their omnivorous diet, ability to stand on two legs, and symbolic importance,[20] and there is widespread support for bear protection, at least in more affluent societies.[21] In more rural and poorer regions attitudes may be more shaped by the dangers posed by bears and the economic costs that they incurr to farmers and ranchers.[19] Some populated areas with bear populations have also outlawed the feeding of bears, including allowing them access to garbage or other food waste. Bears in captivity have been trained to dance, box, or ride bicycles; however, this use of the animals became controversial in the late 20th century. Bears were kept for baiting in Europe at least since the 16th century.

Bears as food and medicine

Many people enjoy hunting bears and eating them. Their meat is dark and stringy, like a tough cut of beef. In Cantonese cuisine, bear paws are considered a delicacy. The peoples of China, Japan, and Korea use bears' body parts and secretions (notably their gallbladders and bile) as part of traditional Chinese medicine. It is believed more than 12,000 bile bears are kept on farms, farmed for their bile, in China, Vietnam and South Korea.[22] Bear meat must be cooked thoroughly as it can often be infected with trichinellosis.[23][24][25]

Culture

Names

The English word "bear" comes from Old English bera and belongs to a family of names for the bear in Germanic languages, in origin from an adjective meaning "brown".[26] In Scandinavia the word for bear is björn (or bjørn), and is a relatively common given name for males. The use of this name is ancient and has been found mentioned in several runestone inscriptions.[27] In Germanic culture, the bear was a symbol of the warrior, as evident from the Old English term beorn which can take the meaning of both "bear" and "warrior".

The reconstructed Proto-Indo-European name of the bear is *h₂ŕ̥tḱos, whence Sanskrit r̥kṣa, Avestan arša, Greek ἄρκτος, Latin ursus, Welsh arth (whence perhaps the given name "Arthur"). Also compared is Hittite ḫartagga-, the name of a monster or predator.[28] In the binomial name of the brown bear, Ursus arctos, Linné simply combined the Latin and Greek names. The female first name "Ursula", originally derived from a Christian saint's name and common in English- and German-speaking countries, means "little she-bear" (dimunitive of Latin ursa). In Switzerland the male first name "Urs" is especially popular.

In Russian and other Slavic languages, the word for bear, "Medved" (медведь), and variants or derivatives such as Medvedev are common surnames.

The Irish family name "McMahon" means "Son of Bear" in Irish.

In East European Jewish communities, the name "Ber" (בער) — Yiddish cognate of "Bear" — has been attested as a common male first name, at least since the 18th century, and was among others the name of several prominent Rabbis. The Yiddish "Ber" is still in use among Orthodox Jewish communities in Israel, the US and other countries.

With the transition from Yiddish to Hebrew under the influence of Zionism, the Hebrew word for "bear", "Dov" (דב), was taken up in contemporary Israel and is at present among the commonly used male first names in that country.

"Ten Bears" (Paruasemana) was the name of a well-known 19th Century chieftain among the Comanche. Also among other Native American tribes, bear-related names are attested.

Myth and legend

"En uheldig bjørnejakt" (An Unfortunate Bear Hunt) by Theodor Kittelsen.
Onikuma from Ehon Hyaku Monogatari
According to his hagiography, a bear killed Saint Corbinian's pack horse on the way to Rome and so the saint commanded it to carry his load. Once he arrived in Rome, however, he let the bear go.

Some evidence has been brought to light on prehistoric bear worship, see Arctic, Arcturus, Great Bear, Berserker, Kalevala. Anthropologists such as Joseph Campbell have regarded this as a common feature in most of the fishing and hunting-tribes. The prehistoric Finns, along with most Finno-Ugric peoples, considered the bear as the spirit of one's forefathers. This is why the bear (karhu) was a greatly respected animal, with several euphemistic names (such as otso, mesikämmen and kontio). The bear is the national animal of Finland.

This kind of attitude is reflected in the traditional Russian fairy tale "Morozko", whose arrogant protagonist Ivan tries to kill a mother bear and her cubs — and is punished and humbled by having his own head turned magically into a bear's head and being subsequently shunned by human society.

"The Brown Bear of Norway" is a Scottish fairy tale telling the adventures of a girl who married a prince magically turned into a bear, and who managed to get him back into a human form by the force of her love and after many trials and difficulties.

There has been evidence about early bear worship in China and among the Ainu culture as well (see Iomante). Korean people in their mythology identify the bear as their ancestor and symbolic animal. According to the Korean legend, a god imposed a difficult test on a she-bear, and when she passed it the god turned her into a woman and married her.

In addition, the Proto-Indo-European word for bear, *h₂ŕ̥tḱos (ancestral to the Greek arktos, Latin ursus, Welsh arth (cf. Arthur), Albanian ari, Armenian arj, Sanskrit ṛkṣa, Hittite ḫartagga) seems to have been subject to taboo deformation or replacement in some languages (as was the word for wolf, wlkwos), resulting in the use of numerous unrelated words with meanings like "brown one" (English bruin) and "honey-eater" (Slavic medved).[29] Thus some Indo-European language groups do not share the same PIE root. The theory of the bear taboo is taught to almost all beginning students of Indo-European and historical linguistics; the putative original PIE word for bear is itself descriptive, because a cognate word in Sanskrit is rakṣas, meaning "harm, injury".[30]

The saddled "bear of St. Corbinian" the emblem of Freising, here incorporated in the arms of Pope Benedict XVI

Legends of saints taming bears are common in the Alpine zone. In the arms of the bishopric of Freising (see illustration) the bear is the dangerous totem animal tamed by St. Corbinian and made to carry his civilised baggage over the mountains. A bear also features prominently in the legend of St. Romedius, who is also said to have tamed one of these animals and had the same bear carry him from his hermitage in the mountains to the city of Trento.

Coat of Arms of the Abbey of Saint Gall

Similar stories are told of Saint Gall and Saint Columbanus.

This recurrent motif was used by the Church as a symbol of the victory of Christianity over Paganism, represented by the fiery.[31]

"The Three Bears", Arthur Rackham's illustration to English Fairy Tales, by Flora Annie Steel

Imaginary bears are a popular feature of many children's stories including Goldilocks and the Three Bears, the Berenstein Bears, and Winnie the Pooh.

The constellations Ursa Major and Ursa Minor represent bears.

Symbolic use

The British Lion, the Persian Cat and the Russian Bear (see The Great Game)

The Russian bear is a common National personification for Russia (as well as the Soviet Union) and even Germany. The brown bear is Finland's national animal.

In the United States, the black bear is the state animal of Louisiana, New Mexico, and West Virginia; the grizzly bear is the state animal of both Montana and California.

Bears appear in the canting arms of Berne and Berlin.

Also, "bear", "bruin", or specific types of bears are popular nicknames or mascots, e.g. for sports teams (Chicago Bears, California Golden Bears, Boston Bruins); and a bear cub called Misha was mascot of the 1980 Summer Olympics in Moscow, USSR.

Smokey Bear

Smokey Bear has become a part of American culture since his introduction in 1944. Known to almost all Americans, he and his message, "Only You Can Prevent Forest Fires" (updated in 2001 to "Only You Can Prevent Wildfires") has been a symbol of preserving woodlands.[32] Smokey wears a hat similar to one worn by many U.S. state police officers, giving rise to the CB slang "bear" or "Smokey" for the highway patrol.

Figures of speech

The physical attributes and behaviours of bears are commonly used in figures of speech in English.

  • In the stock market, a bear market is a period of declining prices. Pessimistic forecasting or negative activity is said to be bearish (due to the stereotypical posture of bears looking downwards), and one who expresses bearish sentiment is a bear. Its opposite is a bull market, and bullish sentiment from bulls.
  • In gay slang, the term "bear" refers to male individuals who possess physical attributes much like a bear, such as a heavy build, abundant body hair, and commonly facial hair.
  • A bear hug is typically a tight hug that involves wrapping one's arms around another person, often leaving that person's arms immobile. It was used in the Ronald Reagan political ad "Bear in the woods."
  • Bear tracking - in the old Western states of the U.S. and to this day in the former Dakota Territory, the expression, "You ain't just a bear trackin'.", is used to mean "You ain't lying" or "That's for sure" or "You're not just blowing smoke". This expression evolved as an outgrowth of the experience pioneer hunters and mountainmen had when tracking bear. Bears often lay down false tracks and are notorious for doubling back on anything tracking them. If you are not following bear tracks, you are not following false trails or leads in your thoughts, words or deeds.
  • In Korean culture a person is referred to as being "like a bear" when they are stubborn or not sensitive to what is happening around their surroundings. Used as a phrase to call a person "stubborn bear."
  • The Bible compares King David's "bitter warriors", who fight with such fury that they could overcome many times their number of opponents, with "a bear robbed of her whelps in the field" (2 Samuel 17:8 [3]). The term "a bereaved bear" (דב שכול), derived from this Biblical source, is still used in the literary Hebrew of contemporary Israel.

Teddy bears

Around the world, many children have stuffed toys in the form of bears.

See also

References

  1. ^ "Slovakia warns of tipsy bears". http://www.earthtimes.org/articles/show/241001,slovakia-warns-of-tipsy-bears.html. Retrieved 2008-11-11. 
  2. ^ Kemp, T.S. (2005). The Origin and Evolution of Mammals. Oxford University Press. ISBN 0198507607. 
  3. ^ Wang, Xiaoming, Malcolm C. McKenna, and Demberelyin Dashzeveg (2005). "Amphicticeps and Amphicynodon (Arctoidea, Carnivora) from Hsanda Gol Formation, Central Mongolia and Phylogeny of Basal Arctoids with Comments on Zoogeography" ([dead link]). American Museum Novitates (3483): 57. http://www.nhm.org/expeditions/rrc/wang/documents/Wangetal2005ShandGolarctoids.pdf. 
  4. ^ Wang Banyue and Qiu Zhanxiang (2005). "Notes on Early Oligocene Ursids (Carnivora, Mammalia) from Saint Jacques, Nei Mongol, China" (PDF). Bulletin of the American Museum of Natural History 279 (279): 116–124. doi:10.1206/0003-0090(2003)279<0116:C>2.0.CO;2. http://digitallibrary.amnh.org/dspace/bitstream/2246/447/22/B279a05.pdf. 
  5. ^ Qiu Zhanxiang (2003). "Dispersals of Neogene Carnivorans between Asia and North America" (PDF). Bulletin American Museum of Natural History 279 (279): 18–31. doi:10.1206/0003-0090(2003)279<0018:C>2.0.CO;2. http://digitallibrary.amnh.org/dspace/bitstream/2246/447/25/B279a02.pdf. 
  6. ^ Lindburg, Donald G. (2004). Giant Pandas: Biology and Conservation, pp. 7–9. University of California Press
  7. ^ Olaf R. P. Bininda-Emonds. "Phylogenetic Position of the Giant Panda". In Lindburg, Donald G. (2004) Giant Pandas: Biology and Conservation, pp. 11–35. University of California Press
  8. ^ The Brown Bear: Father of the Polar Bear?, Alaska Science Forum
  9. ^ a b c d e f Garshelis, David L. (2009). "Family Ursidae (Bears)". in Wilson, Don; Mittermeier, Russell. Handbook of the Mammals of the World. Volume 1: Carnivores. Barcelona: Lynx Edicions. ISBN 978-84-96553-49-1. 
  10. ^ Bunnell, Fred (1984). Macdonald, D.. ed. The Encyclopedia of Mammals. New York: Facts on File. pp. 87. ISBN 0-87196-871-1. 
  11. ^ Mattson, David J. (1998). "Diet and Morphology of Extant and Recently Extinct Northern Bears". Ursus, A Selection of Papers from the Tenth International Conference on Bear Research and Management, Fairbanks, Alaska, July 1995, and Mora, Sweden, September 1995 10: 479–496. http://www.jstor.org/stable/3873160. 
  12. ^ Ryan, Christopher; Pack, James C.; Igo, William K.; Billings, Anthony (2007). "Influence of mast production on black bear non-hunting mortalities in West Virginia". Ursus 18 (1): 46–53. doi:10.2192/1537-6176(2007)18[46:IOMPOB]2.0.CO;2. 
  13. ^ Matheus, Paul E. (1995). "Diet and Co-ecology of Pleistocene Short-Faced Bears and Brown Bears in Eastern Beringia". Quaternary Research 44 (3): 447–453. doi:10.1006/qres.1995.1090. 
  14. ^ Joshi, Anup (1997). "Seasonal and Habitat-Related Diets of Sloth Bears in Nepal". Journal of Mammalology 1978 (2): 584–597. 
  15. ^ a b Hwang, Mei-Hsiu (2002). "Diets of Asiatic Black Bears in Taiwan, with Methodological and Geographical Comparisons". Ursus 13: 111–125. 
  16. ^ Zager, Peter; Beecham, John (2006). "The role of American black bears and brown bears as predators on ungulates in North America". Ursus 17 (2): 95–108. doi:10.2192/1537-6176(2006)17[95:TROABB]2.0.CO;2. 
  17. ^ Clark, Douglas (2003). "Polar Bear-Human Interactions in Canadian National Parks, 1986-2000". Ursus 14 (1): 65–71. 
  18. ^ a b Fredriksson, Gabriella (2005). "Human–sun bear conflicts in East Kalimantan, Indonesian Borneo". Ursus 16 (1): 130–137. doi:10.2192/1537-6176(2005)016[0130:HBCIEK]2.0.CO;2. 
  19. ^ a b c Goldstein, Issac; Paisley, Susanna; Wallace, Robert; Jorgenson, Jeffrey P.; Cuesta, Francisco; Castellanos, Armando (2006). "Andean bear–livestock conflicts: a review". Ursus 17 (1): 8–15. doi:10.2192/1537-6176(2006)17[8:ABCAR]2.0.CO;2. 
  20. ^ Kellert, Stephen (1994). "Public Attitudes toward Bears and Their Conservation". Bears: Their Biology and Management 9 (1): 43–50. doi:10.2307/3872683. 
  21. ^ Andersone, Žanete; Ozolinš, Jānis (2004). "Public perception of large carnivores in Latvia". Ursus 15 (2): 181–187. doi:10.2192/1537-6176(2004)015<0181:PPOLCI>2.0.CO;2. 
  22. ^ "BBC Test kit targets cruel bear trade". 2007-06-11. http://news.bbc.co.uk/2/hi/science/nature/6742671.stm. Retrieved 2010-01-01. 
  23. ^ "Trichinellosis Associated with Bear Meat". http://www.cdc.gov/MMWR/preview/mmwrhtml/mm5327a2.htm. Retrieved 2006-10-04. 
  24. ^ "BBC News - Bear meat poisoning in Siberia". 1997-12-21. http://news.bbc.co.uk/1/hi/world/europe/41529.stm. Retrieved 2006-10-04. 
  25. ^ "Finnish Food Safety Authority: Bear meat must be inspected before serving in restaurants". http://www.evira.fi/portal/en/food/current_issues/?id=346. Retrieved 2006-10-04. 
  26. ^ Pokorny (1959) [1]
  27. ^ http://hildebrand.raa.se/arkeologi/uppland.asp
  28. ^ Pokorny (1959) [2]
  29. ^ Votruba, Martin. "Bears". Slovak Studies Program. University of Pittsburgh. http://www.pitt.edu/~votruba/qsonhist/bear.html. Retrieved 2009-03-12. 
  30. ^ The Brown One, The Honey Eater, The Shaggy Coat, The Destroyer
  31. ^ Michel Pastoreau (2007)L'ours. Historie d'un roi déchu
  32. ^ Ad Council : Forest Fire Prevention - Smokey Bear (1944–Present)

Further reading

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  • Bears of the World, Terry Domico, Photographs by Terry Domico and Mark Newman, Facts on File, Inc, 1988, hardcover, ISBN 0-8160-1536-8
  • The Bear by William Faulkner
  • Brunner, Bernd: Bears — A Brief History. New Haven and London: Yale University Press, 2007

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How do your get bearings out?
What is the bearing?

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